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Introduction Follwing the studies on movement, activity, home range and habitat use of red deer in the Bavarian
Forest National Park during the last years (see below) seven red deer were fitted with GPS GSM collars in spring 2004. The investigation area is located in the south-east of Germany (figure 1).
Fig. 1: Location of the investigation area in Germany (red symbol)
A part of the red deer population in the Bavarian Forest National Park overwinters in 3 “winter enclosures”. This
red deer management concept greatly helped reduce browsing and stripping damage to trees and saplings. This
winter enclosure management method has been proved successful since the 1970’ and `80`. A total of 80 red
deer spend the winter in this enclosure and are released in the spring usually during the first weeks of May 2004.
The GPS receivers are programmed to take fixings every 2-4 hours and to send the data by SMS immediately after 7 fixings are taken via the mobile phone system directly to our groundstation at the office. The animals were
caught in a corral and fitted with GPS-GSM collars (figure 2 and 3).
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Fig. 2: A young stag in a corral
Photos: Marco Heurich, Rainer Pohlmann
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Three of the collars were already used last year and equipped with new batteries. All collars are state of the art:
If a collar is out of GSM coverage the GPS fix data will be immediately transmitted when GSM coverage is available. This is the most secure option to get the maximum of GPS data via GSM network.
Tab. 1: Data of the collared red deer, 2004
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Collar ID 2004
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Name
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Day of collaring 2004
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Sex
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Age
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694
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Alois
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April 16
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male
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2
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696
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Benedikt
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May 2
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male
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6
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841
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Fritz
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April 27
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male
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3
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449
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Willi
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April 7
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male
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9
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878
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Oma
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May 5
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female
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12
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877
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Paul
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April 27
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male
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8
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879
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Susi
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May 5
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female
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2
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The red deer were released from the enclosure on May, 2004. The movements are presented in figure 4. The maps are updated all 8 weeks.

Fig. 4: Location points of 6 red deer (legend: age in bracket; n=8678 fixes) connected in chronological order, after
leaving the winter enclosure, May 13 - December 1, 2004 ©Basisdaten der Nationalparkverwaltung Bayerischer Wald
Introduction Follwing the studies on movement of red deer in the Bavarian Forest National Park during the last years (see below) eight red deer were fitted with GPS-GSM collars in spring 2003.
The GPS receivers are programmed to take fixings every 2-4 hours and to send the data by SMS immediately after 7 fixings are taken via the mobile phone system directly to our groundstation at the office. The animals were caught in a corral and fitted with GPS-GSM collars (figure 2 and 3).
Tab. 1: Data of the collared red deer, 2003
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Collar ID
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Name
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Day of collaring 2003
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Sex
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Age
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448
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Kasimir
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April 8
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male
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11
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306
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Bine
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April 30
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female
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14
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452
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Oma
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March 18
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female
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11
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450
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Spitz
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March 24
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male
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3
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449
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Cäsar
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April 8
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male
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7
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453
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Schwalbe
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March 18
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female
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2
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451
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Fritz
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March 19
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male
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2
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302
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Charly
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April 30
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male
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2
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The red deer were released from the enclosure on May 13, 2003. The movements are presented in figure 4 for
males and 5 for females. It is obviously, that the male red deer show a more complex space behaviour and cover larger home ranges as the females.

Fig. 4: Location points of 3 male red deer connected in chronological order, after leaving the winter enclosure, May 13 - October 6, 2003
©Basisdaten der Nationalparkverwaltung Bayerischer Wald

Fig. 5: Location points of 3 female red deer connected in chronological order, after leaving the winter
enclosure, May 13 - October 6, 2003 (n= 959, 1442 and 1229)
The activity of the tracked red deer has been monitored by a two axis activity logger. By this, the acceleration of
the collar will be estimated 6-8 times every second and the overall mean value has been stored every 5 minutes.
The complete GPS data sets and the activity and temperature data sets from the tracking period 2003 have been
stored on board of the GPS GSM collar and were outried by computer via Link Manager. The activity of hint 452 are shown in figure 6.
Fig. 6: Activity data of female 452 in x- and y direction (blue and green symbols) and temperature data (red line), April 30 2003 to March 24, 2004
Study 2002
Investigations on the use of habitat of red deer in the Bavarian forest are part of a cooperative pilot project
between the Management of the Bavarian Forest National Park and Environmental Studies. The GPS collars are constructed by Vectronic Aerospace, Berlin. The company has financed a part of the project costs.
The Project In March and April 2002 three red deer (female, young stag and older stag) in a winter enclosure were caught
and provided with a GPS-Plus GSM collar. A total of 80 red deer spend the winter in this enclosure and are
released in the spring usually during the first weeks of May. The red deer were outfitted with collars prior to
release to test the data transmission system. The collars are programmed to take fixings every 3 hours and send
the data by SMS on a daily basis. The red deer were released from the enclosure about the May 8, 2002. Their movements until December 2, 2002 are presented in figure 2.
Fig. 2: Location points of 3 red deer connected in chronological order, after leaving the winter enclosure, May 8, 2002 - January 4, 2003
©Basisdaten der Nationalparkverwaltung Bayerischer Wald
The young stag spent most of the time in the higher mountain areas on both sides of the border between
Bavaria and Czech Republic. In this area the GSM coverage is poor, that is why the data transmission is insufficient.
In August the female red deer and the stag spent their time in higher regions, probably because of the
disturbance by mushroom seekers in the lower areas. On September 7, the stag moved across the border to the
Czech Republic, expanding his home range. From September 24 to October 23 no data was transmitted. Probably
the animal lived in an area without GSM coverage. Since October 23 the data arrived as usual. The stag moved back to the border.

Fig. 3: Location points of the 3 red deer from December 1, 2002 - January 4, 2003 ©Basisdaten der Nationalparkverwaltung Bayerischer Wald
In the last two weeks of September the female red deer moved down from the higher mountain areas and stayed
in the first weeks of October in or nearby the winter enclosure. Whereas the young stag moved at this time into
the higher mountain regions and expands his home range into Czech territory, travelled 9 km to the east, visited the winter enclosure and travelled back to the higher mountain regions.
At the time the female red deer are dwelling in the winter-enclosure with 65 others, the two stags are still
moving in higher mountain regions, close to the border of the Czech Republic. This is unusual behaviour, because
normally red deer pass the winter in the winter-enclosure or stays in the lower foothills. But this winter, until the
first week of January, the mountains were nearly free of snow and obviously food was still available.
Statistics from May 8 - December 2, 8 fixings/day
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Collar statistics
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female
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stag
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young stag
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fixings possible
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1666
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1666
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1666
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fixings via SMS transmitted
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1459
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1067
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951
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fixing/SMS efficiency [%]
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88
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64
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57
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red deer statistics
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max. distance between 2 positions [m]
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4377
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8993
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8211
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mean distance of all positions [m]
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299
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320
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469
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home range MCP [ha]
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1214
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4726
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6131
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home range Kernel (95%) [ha]
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616
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426
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1389
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Whereas the collar of the female red deer has a good efficiency (88%), the data transmission of the stag’s collars
is less (57% and 64%). It depends on the bad GSM coverage in the border area of the Czech Republic, where the stags spend a lot of time in the higher mountain regions.
Results from an Investigation with “normal” GPS-collars on red deer: Study 2001
In April 2001 5 female red deer from two different winter enclosures in the Bavarian Forest National Park were
outfitted with GPS-Plus collars. At present one collar has been retrieved by recapturing one of the animals. The
collar contained recordings of 6500 localizations from April 2001 to March 2002, whereby positions were recordedhourly. The migratory paths of the animal are shown in Fig. 5.

Fig. 5: Location points of a female red deer connected in chronological order, May 2001 - March 2002 (n = 6500)
The 3 year old female red deer showed a conspicuous migratory pattern. It only wandered to the periphery of its
range from the central area to which it quickly returned. Longer treks or excursions over several days outside of
its home range did not occur.The animal was very homebound in choosing its winter quarters. It spent the winter in the same winter enclosure as the previous year.
The female red deer first remained in the vicinity of this enclosure upon its release on May 15, 2001. Later it
moved to an area 5 km to the north. The animal often moved back and forth through a small corridor between these two ranges. The upper range encompassed an area of 843 ha, the lower an area of 900 ha (calculated by
the convex polygon method, see also first introduction to our work with satellite telemetry).
More information about the project
The investigations should provide data to aid in the decision making process in the management of the Bavarian Forest National Park.
After the establishment of the Bavarian Forest as a national park attempts were made to develop solutions for
the management of red deer in such a setting. On the one hand the high population density of the red deer was
reduced and on the other unfenced feeding stations were removed in favour of 3 enclosed ones for the winter
(“winter enclosures”), where the animals could overwinter. This management concept greatly helped reduce browsing and stripping damage to trees and saplings. This winter enclosure management method has been
proved successful since the 1970’ and `80`.
An investigation conducted in 1984 showed that extensive areas in the Bavarian Forest National Park were not
used by the red deer. The majority of the red deer in this park overwintered in the enclosures and had their summer ranges on Czech territory between the national border and the farther border fence. In the past few
years several changes have occurred to influence the habitat and behaviour of the red deer:
- The removal of the border fence has permitted a greater movement of individuals between the Bavarian Forest National
park and the Sumava National Park. Especially a migration of the red deer from the Bohemian Forest to the unoccupied areas in the Bavarian Forest is assumed.
- Due to the extemsive clear cuts on the Czech side and the large contiguous areas of deadwood in the Bavarian Forest,
the browse conditions for the red deer have substantially improved. Helicopter logging and the lack ofcover apparently cause achange in habitat use among the red deer.
- The scant snowcover during the past few years and the consequent greater availability of browse and increased mobility
permitted the red deer to even overwinter on the upper slopes.
- The recent occurrence of wild boar and lynx in this area could also affect a change in the temporal-spatial behavioural
system of the red deer.
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Fig. 4: View from the village of St. Oswald towards the Rachel.Behind the mountain range lies the National Park Sumava in the Czech Republic.
The red deer management system effectively practiced for years must now be adapted to the changed
conditions.For this purpose, an exact analysis of the above mentioned changes and an evaluation of alternative
solutions is necessary. The present investigation is the first step in obtaining basic data on the actual use of
habitat by red deer in the Bavarian Forest National Park and in the Sumava National Park.The following questions must be answered:
- What kind of migratory behaviour do the red deer actually show?
- Are there at present still potential overwintering areas? And if so, can the animals reach them and
what habitat condtions prevail there?
- How would the animals react if the winter enclosures were removed, and what would the consequent management strategy be?
Author's e-mail address:
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